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had been colonizing

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He explained how Candida albicans, an aggressive sugar-eating yeast that had been colonizing my intestines, is a common concern for celiacs, whose tiny, hair-like villi in their intestines have been flattened and damaged by gluten.

From New York Times Jul. 20, 2010

France, too, had been colonizing in America, and had drawn her frontier line from the mouth of the Mississippi to Canada.

From A Short History of Spain by Parmele, Mary Platt